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However, the dimensions of the heterogeneities of plankton biomass and bioluminescence intensity can differ, as was noted above.然而，浮游生物量非均质性和发光强度的尺寸可以不同，正如上文所述。 This might be due to the fact that only a section of the plankton这可能是由于这样的事实，只有一节的浮游生物community bioluminesce, and it is this part which is associated with diel rhythms in bioluminescence in the studied regions (Bityukov et al., 1993).社区生物发光，这是这是与生物发光昼夜节律在所研究的地区（比秋科夫等相关部分。，1993）。The increase of spatial heterogeneities at night could be based upon vertical migration and the behavioural mechanisms of the planktonic organisms forming the patches.空间异质性的增加可能会在夜间基于垂直迁移和形成的补丁浮游生物的行为机制。 Patches of organisms can be maintained by visual contacts, chemoreception and vibroreception生物体的补丁可维持视觉接触，化学感受和vibroreceptionbetween and among species (Newbury,1972; Omori and Hamner, 1982).间和种间（纽伯里，1972;大森和哈姆纳，1982）。Our observation that the number of spectral peaks of the bioluminescent, plankton and acoustic fields change during the day cycle, on a background of more stable statistical structure of the temperature field heterogeneity, also supports the idea that biological mechanisms can be powerful at certain times (at night我们的看法，即白天周期数项发光，浮游生物和改变声场谱峰在一个更稳定的温度场结构异质性的统计背景，也支持生物机制的想法，可以在某些时候强大（夜间for instance) in the formation of these heterogeneites.例如）在这些heterogeneites形成。 The biological mechanisms acting on the meso and microscales should interact with the other biological factors and mechanisms acting on larger scales.关于中观和微量的行动应与其他生物因素和机制的作用较大规模的生物机制。 In the upper layers of the Black Sea the fluctuations in the diel rhythm of zooplankton abundance can be affected by the synchronised rhythms of predator/prey在黑海中的浮游动物丰度昼夜节律的波动可以通过捕食同步的节奏影响/猎物上层interactions and the reproduction rate of zooplankton populations (Kovalev, 1993; Zagorodnaya and Kovalev,1988).相互作用和浮游动物种群（科瓦廖夫，1993年繁殖率; Zagorodnaya和科瓦廖夫，1988年）。 In general, zooplankton reproduction rate acts as an ecologically important factor in the formation of spatial heterogeneity in the microscale在一般情况下，浮游动物繁殖作为一个在空间异质性形成的重要生态因子率行为的微观(Owen, 1989). （欧文，1989年）。On the other hand, the general agreement between the bioluminescence, plankton and temperature spectral functions indirectly indicate that spatial-temporal variability of these fields can be influenced by other factors causing this similarity.另一方面，生物发光之间的广泛协议，浮游生物和温度谱函数间接表明，时空变化，这些领域可引起此相似的其他因素的影响。 Previous field studies of microscale variability of zooplankton abundance together with turbulent diffusion have indicated good微型浮游动物的丰度与变异性的湍流扩散以及前场的研究表明良好agreement of spectral functions with the concentration of tracer dye and species abundance in the upper layers of the Black Sea (Nemirovski et al., 1991).协议谱函数与示踪染料和物种丰富度在黑海（上层浓度涅米罗夫斯基等人。，1991）。A number of maxima in the spectral analysis of the SVBS distribution in the surface layer remained constant throughout the 24 hr period (as shown in Table 1).一个极大值在表面层中的分布SVBS光谱分析人数仍然在24小时内（如表1所示常量）。 A level of SVBS at 80 kHz frequency in the Black Sea is connected with the abundance of一SVBS在80千赫频率黑海连接水平与丰富的macroplankton populations (Medusae, Ctenophora) and fish larvae which do not extensively migrate out of the upper isothermal layer in daytime (Tokarev & Sokolov, 1993). macroplankton人群（水母，Ctenophora）和鱼类幼虫不广泛迁移出白天等温层上部（托卡列夫和索科洛夫，1993）。 There is a considerable increase (in2–4 times) in sound reverberation in patch sizes at有一个相当大的增长（平方英寸- 4倍）在补丁大小声混响在night, which can be connected (besides the increase in zooplankton abundance) with the change in spatial orientation of organisms.夜晚，它除了可以连接在浮游动物数量增加（）与在空间定位改变的生物体。